The group “Testudines” usually represents the turtles, tortoises, and terrapins. Recentstudies on Testudines indicate turtles are rich in both α- and β-keratins (Dalla Valleet al. 2013). The β-keratin/α-keratin ratio is higher in the carapace in hard-shelledturtle. This ratio provides an important factor for further study in the background ofkeratin.